Allies become enemies when both Casey and Frannie are nominated for the title of Omega Chi Sweetheart. Who will prevail in this the ultimate cat fight? Meanwhile, as Rusty's relationship with Jen K. heats up, he longs for more private time with her but his judgmental roommate Dale is always around, making it difficult for the two lovebirds to spend time alone.

When a scandalous story in the college newspaper hits stands trashing the Greek system, the brothers and sisters of Omega Chia, Zeta Beta Zeta and even Kappa Tau are in a frenzy to figure out who the undercover culprit could be. Meanwhile, after navigating some troubled waters with her boyfriend, Ashleigh turns to Calvin for support and even goes so far as to hit on him, only to learn he has a secret.

Hoping to sneak around Dean Bowman's probationary despotism, Casey tries to liven up the Zeta Betas by joining forces with Cappie and his Kappa Tau brothers to plan a Prohibition party with a Great Gatsby theme.

Trouble is caused by Casey when she bends the sorority house rules for Ashleigh. Cappie tries to get rid of restrictions.

Meanwhile, Rusty finds himself attracted to Tina, the girl who wants to take down the Greek system.

Rusty attempts to solve a mystery involving a secret service he ran across on campus in hopes of impressing Cappie. Meanwhile, some of Frannie's defectors from ZBZ house have asked to return, and Rebecca seeks dating advice from Calvin as she readies to go out with Robin.

A lonely Casey meddles in Rusty's new romance with Jordan. Elsewhere, Calvin finally gets a new roommate, and then he discovers that the guy has a surprising secret; and Evan and Cappie reignite their rivalry, this time over a secret-society member.

The season opens with Casey, Evan, & Ashleigh’s graduation day. Casey and Cappie haven’t spoken since Spring Break and are anticipating saying their final goodbyes to each other, which does not go as smoothly as expected. We then skip to the end of summer and Casey packing up to move to Washington DC for law school. Through some prompting from Evan, Casey learns that the reason she did not get into CRU law school was a result of a bad recommendation from Joel (Sam Page). After Casey confronts Sam, he reconsiders and has his letter rescinded. Casey is accepted into CRU law and decides to stay after all. Ashleigh has moved to New York for an internship that is not exactly what she was hoping for. Evan is attending CRU law school with Casey.