When Casey discovers that Cappie is taking the same women’s studies class taught by Professor Freeman, she knows that it will complicate her relationship with Max. As a result, Casey decides to enlist the help of her new professor to help Max and Cappie get along. Meanwhile, Rusty starts his first elective class in Art History, and as a lucky bonus meets a pretty new transfer student, Jordan. Although he’s nervous about talking to her, he finds the courage to invite her to a KT party. But the night of the party he learns that some guys have all the luck, namely the new KT pledge, Andy. At the ZBZ house, Ashleigh hires a new hasher, Fisher, who draws a lot of attention from the sisters of ZBZ but, Fisher only has eyes for Ashleigh.

Troubled by the recent discovery that Andy hooked up with Jordan, Rusty is hesitant to take Andy on as his little brother. Will Rusty be able to get past the budding relationship to become the big brother Cappie has been to him? When Cappie hears of Rusty’s reluctance, he takes Rusty aside to remind him that he brought Andy as a pledge to the fraternity and it is his responsibility to be the best Big Brother that Rusty can be for him. Meanwhile, when Evan’s real-life brother, Patrick, comes for a visit, their celebration at Dobler’s is interrupted by Cappie and the three end up reminiscing about their old camp days. But later Patrick shocks and disappoints Evan by asking for money. Frustrated by his brother’s actions, Evan takes it out on Frannie. When Casey and Rebecca believe that the thief responsible for stealing the ZBZ food supply is Fisher, they can’t figure out why Ashleigh won’t accept it. But when they dig a little further, Ashleigh reveals the delicious truth.

When Frannie challenges the sisterhood of ZBZ girls, and Casey retaliates by challenging the IKI’s to a game of dodgeball. Meanwhile, Rusty struggles with the fact that Dale is so in love with their new apartment that he never leaves, so Rusty enlists the Kappa Tau’s to invade the house to scare him away. But the plan backfires when the landlord shows up the next morning with a tour.

Casey’s grand-big sister, Kiki , is back at Cyprus-Rhodes University for her wedding and all the ZBZ girls are invited. Amidst all the wedding frenzy, Max decides it’s time to take his relationship with Casey to the next level and gets down on one knee. Meanwhile, with Andy out of town, Jordan ends up dragging Rusty around town to help pick up the flowers for Kiki’s wedding. When the two discover that the order was never placed, they have to come up with a plan of action however, the plan backfires and puts both of them in the emergency room with identical allergic reactions. The unexpected common bond leads them to share a kiss. Rebecca, in the meantime, soon finds herself in new territory as she kisses Ashleigh’s openly gay big sis, Robin, in one of the bachelorette party game sparking Rebecca’s curiosity.

The KTs race to return a stash of stolen campus goods, hoping not to get caught by security. Elsewhere, Casey sneaks Max into a faculty party so he can meet a visiting professor that needs an intern, and Ashleigh considers taking her relationship with Fisher public.

Two girls that Cappie is dating at the same time both end up in his study group. Meanwhile, Rusty is having doubts about his relationship with Jordan, and Rebecca solicits help to retrieve her laptop.

A lonely Casey meddles in Rusty's new romance with Jordan. Elsewhere, Calvin finally gets a new roommate, and then he discovers that the guy has a surprising secret; and Evan and Cappie reignite their rivalry, this time over a secret-society member.

Rusty and Ashleigh try hard to win the float competition, while Casey and Cappie try avoid each other as Homecoming Week heats up. Elsewhere, Evan gets word that he may lose his trust fund, and Ashleigh uncovers one of Fisher's secrets.

After she falls into a manhole risking her life, coupled with the Kappa Tau's throwing an "End of the World" party, Casey is sent into an emotional tailspin, questioning her relationship with Max and her lingering feelings for Cappie. Meanwhile, Ashleigh faces her own punishment with Pan Hellenic after a complaint is lodged by Frannie for the Homecoming Float sabotage. Ashleigh turns to Casey to help fight the charge, but when Casey gets caught up in her own personal drama, she leaves Ashleigh to face Frannie alone. And with Dale having trouble keeping his cougar girlfriend's sexual advances at bay and Calvin trying his best to resist hooking up with his new hot roommate, Grant. Dale convinces Calvin to go in on a "purity pledge" partnership so they can help each resist their most recent temptations.

The students at CRU feel the ramifications of the Kappa Tau's "End of the World" Party. Rusty risks being dropped from the honors program if he cannot raise his grade in organic chemistry and Casey adjusts to single life.

Casey finds out about Rebecca and Fischer's secret kiss, but when Rebecca confesses to Ashleigh, she outs Casey for keeping it a secret. When Mr. Cartwright, Mr. Howard, and Mr. Logan arrive for Father-Daughter weekend at ZBZ, they find out about their daughters' feud, and it reflects upon their weekend behavior. At the same event, Rusty gets nervous about meeting Jordan's father. Meanwhile, in an effort to get over their lost loves, Dale and Cappie fawn over the same girl, but they both ultimately get rejected.

It's time for the annual all-Greek "Undie Run", and the Pan-Hellenic board voted strongly against it. In an effort to change the vote, Casey uses her political savvy to enlist IFC president, Evan, to stage a petition signing. Meanwhile, it's nearing the end of the Greek-wide game of Gotcha!, and only Rusty and Jordan are left, which puts a damper in their effort to spend alone time together. Additionally, Evan finds out about Calvin and Grant's relationship and helps them keep it a secret from the rest of Omega Chi.

Shocked to learn that ZBZ has slipped in the ranks down to fourth, Casey is determined to bring ZBZ back to the number one spot by motivating their pledge class to take action, believing that, if they create a cool photo shoot for the Lamda Sig calendar, it could improve their house’s ranking. However new pldge Abby convinces the other pledges to boycott the shoot so, in order to make nice with the ZBZ pledges, Ashley and Casey get them all makeovers. Meanwhile, dissatisfied with her college experience, Jordan wonders how to make her time at CRU more fulfilling and, in an effort to prove his love and commitment for Jordan Rusty decides to lavaliere her. When Rusty divulges his plans to his KT brothers, they warn him of the Kappa Tau curse that occurs when a brother lavalieres his girlfriend. Elsewhere, Cappie helps Evan find a part time job to help his financial situation, but when he meets one of Evan’s new and very attractive co-workers, Lana, he decides to get a job too.

It's Founders Weekend at the ZBZ House and Casey uses the opportunity to scrap for donations from the alums while Rusty tries to keep Jordan from dropping out of school.

Casey's detective skills inadvertantly start a brawl between the Kappa Tau's and the Omega Chi's. Meanwhile, Rusty, Calvin, and Dale take a trip to Comic-Con as Ashleigh and Fisher resume their relationship but still have a few issues.