Harry vows revenge by taking down the head of Russia's intelligence service in Britain with the help of Lucas North. Lucas's loyalties, however, are secretly no longer with MI5 and he exploits Harry's trust to help a Russian submarine destroy Britain's communications system, causing financial and social chaos.

With Lucas back within MI5 he reveals that the Russians interrogated him about Operation Sugar Horse, the most secret ongoing spy operation MI5 ever had. If the Russians know about it, then MI5 has a very high level mole. Harry seeks out his mentor, The Spycatcher, to help him find the mole. Ben is undercover with an Al-Qaeda cell; they are planning to bomb London and what looks like a dry run unexpectedly turns live.

Harry is imprisoned and accused by MI5 of being the Russian mole. Lucas disappears to Moscow in a bid to gather intelligence and get to the bottom of Harry's arrest, whilst the rest of the team tries to prove his innocence on the Grid. But with so much evidence against him and a traitor among the team, things aren't looking good for Harry.