Out of nowhere, Shiroe finds himself in the city of Akihabara, inside the game "Elder Tale". Thirty thousand Japanese players have been locked inside the game! The world on the other side of the screen becomes reality, and amidst the chaos Shiroe finds his old friend Naotsugu. And as they relax at finding one another, they meet the powerful assassin "Akatsuki". Akatsuki has a request to make of Shiroe...

Serara, a young female member of the guild "Crescent Moon Alliance", is trapped by mysterious men in the northern city of Susukino and needs help. Shiroe and his friends travel to reach her. Annoyed by the game's tasteless food, they travel northwards, and go inside "the depths of Palm", a crumbling series of old mining tunnels. There they meet creepy monsters...

Shiroe and his group take Serara southwards. On the way, they stop at the house of one of the "People of the Land". The people of the land were originally only characters in the game, with no personalities or unique traits of their own, but as Naotsugu talks to them, something seems odd... The party gradually realizes that the world isn't the same as it was when it was a game.

Nyanta told Shiroe the secret of making real food in this world. With Marielle's help, the Crescent Moon Alliance begins a fast food stand, and their "Crescent Burger" is hugely popular with players who are sick of tasteless food. BUt Shiroe's real goal isn't to sell hamburgers, and with Henrietta he begins to work on the next phase of his plan.

To make the city of Akihabara safe, Shiroe calls all the major guild masters to the guild building. His goal is to create a Round Table Conference, which will rule the city. But everyone there wants something different, and his plan is about to go bad. What amazing trick does Shiroe have? Meanwhile, behind the scenes of the conference, the plan to free the younger players from the evil guild Hamelin continues. The strategist SHiroe's quiet battle in Akihabara continues!

Shiroe confronts the 10 guild masters who represent Akihabara with the need to revitalize the town and bring peace and order. His calm and forceful manner brings them over, and the government of Akihabara, the "Round Table", is founded. After this work is done, his guild, "Log Horizon", joyously moves to their new location. That night, Nyanta fixes them his special curry.

The young players travel the Xantleaf peninsula at the camp. Minori, Touya and Serara meet up with new players Isuzu the bard and Rundel Haus the sorcerer. They head into the "La Granda Forest" dungeon, where skeletons await them. The easily-spooked Serara, the strong-willed Touya, and the proud Rundel Haus... will the five of them be able to work together in their first party fight?

Zantleaf is attacked by monsters from both the land and the sea, as the adventurers continue to defend it. Inland, Crusty's army launches a raid on the goblin's main force. At the town of Choushi by the sea, the new players are fighting too.

The second day of the Scales Festival in Akihabara arrives. Akatsuki has gotten closer to Shiroe, and so is very happy all day, and even seems to be enjoying her fights with Naotsugu. Meanwhile, Minori is uncertain with how to deal with her feelings towards him, and goes to speed to Karashin. Karashin is overloaded with paperwork so she decides to help. It seems like a lot of land people merchants have come to Akihabara...

Marielle and Henrietta put on a fashion show as the town reels from the Minami decoy operation. Akatsuki walks down the runway in her colorful clothes, but Shiroe is nowhere to be seen. That night, Shiroe is met by a beautiful and enchanting "queen of darkness." Shiroe is tempted he reaches the heart of the conspiracy...