前往異境的MAR總帥夏赫利亞·連獲得了罪神的遺產,即古代超級武器「眷獸彈頭」,並計劃將世界再次置於天部的統治之下。在這種壓倒性的武力面前,獅子王機關和攻魔局決定極度保密地破壞連接異境和現世的「門」──絃神島。
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雪菜為了思考從上層部發布的無情任務而煩惱。為了保護重要的東西,古城作為絃神島的領主,與同伴一起迎接最後的一戰。
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沙赫爾雅·蓮試圖通過摧毀東京向古城和其他人展示核彈眷獸的力量。與此同時,率領第二次降臨的騎士艦隊的拉·芙利亞襲擊了死都。
古城和雪菜進入異境,阻止沙赫爾雅·蓮並拯救奧蘿拉。
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