Julien, Maurice and Pancho regroup aboard a submarine and ponder their next move, while a captive Mort plots to take down the tyrannical new king.

Clover hopes Sage's guru can help with a plan to oust Koto, who's now made Mort his jester. Julien and crew meet an enchanting tribe of lemur sirens.

As Julien and company head back to Madagascar, their submarine runs into trouble. Mort puts on a play for Koto to cover up the captives' escape.

Julien and the boys team up with sharks and try to form an army. Jarsh-Jarsh's map leads Clover and Sage into danger. Mort's escape plan hits a snag.

Julien and Maurice face a harrowing fight at gladiator school. Mort travels to another dimension. Sage convinces Clover there's still good in Koto.

Julien and Karl set out to stop Koto's wedding to the Crocodile Princess. Clover deals with Koto her own way. Mort fights to bring back "Smart Mort."

As Koto prepares a sacrifice to kick off his wedding, Clover and Sage unleash their new weapon. Mort, Maurice and Pancho arrive with reinforcements.

Nearly vanquished by Koto's troops, Julien rallies Mort's army, the bell snails, the space chimps, Clover and other loyal friends for one last hurrah.