A grisly murder stokes class division on Snowpiercer, the Great Ark Train. Melanie Cavill, the powerful head of hospitality, deputizes a dangerous rebel to help solve the killing -- Andre Layton, the world's only surviving homicide detective.

Layton uses his new position as train detective to investigate the murder while gathering intel for the revolution on the side. Meanwhile, Melanie faces a resource crisis, with potentially drastic consequences for the entire train.

Layton descends into the black market with Till, searching for both the killer and a valuable commodity for his revolution. Melanie stages a prize fight to distract the passengers from mounting class tension.

Revolution has finally come. As Layton leads the lower classes forward in armed rebellion, Melanie's house of cards collapses.

The rebel forces are on their heels when a dangerous foe joins them. A plan is hatched to take the train, but it may destroy them all.