＜RUNWAY＞雜誌是紐約時尚界聖經，總編米蘭達是難纏的大人物。只要稍有自尊的人，絕對無法在她手下工作，可是進入＜RUNWAY＞又是所有對時尚有熱情的年輕女孩最大的夢想。 剛畢業的社會新鮮人安德莉亞（安海瑟薇飾演），憑著西北大學的漂亮履歷，進入＜RUNWAY＞位於曼哈頓總部辦公室。不服輸的她準備擔任米蘭達的助理。這名從頭到腳全是Prada的女士，將為她的人生帶來最殘酷的考驗。 漸漸地，安德莉亞讓自己融入米蘭達要求的氣質和形象，很短時間裡，她學會了這一行說話的模樣身段及行事方法。只是，當她越嫻熟米蘭達所在世界的遊戲規則，她也發現這其實是一個表面上精彩華麗但其實很孤獨的世界，值得嗎？安德莉亞靠自己找到她的答案。

Edward Kaminsky, an aging ad man, wants a golden parachute from his agency; he must first land the Opel auto contract. Rosa, a youth with wealthy parents, wants to establish herself as an artist. The clumsy and enthusiastic Viktor, not quite honest, wants work. When he wanders into Kaminsky's meeting with Opel and says something about irony, the Opel director wants him in on the campaign. Then he steals an idea from Rosa that the Opel director loves. Before Rosa discovers he's expropriated her idea, Rosa and Viktor become lovers. Father-son feelings materialize between Kaminsky and Viktor. Can the impulsive Viktor hold it together before Rosa learns the truth and flies away?