革命軍參謀長薩博聯繫了無尾熊和卡瑪巴卡王國的所有人。他向他們解釋了自己被指控的事情，但偷聽電話的人採取了驚人的行動。

魯夫和喬巴與神秘女孩一起掉進了波濤洶湧的寒冷大海。神兵想要幫助他們，但未知生物又發動了攻擊。同時，桑尼號被魚雷擊中，草帽團的其他成員也被丟進了大海。

從大機器人出來的小女孩給自己取名為維加龐克博士，這讓法蘭奇和其他草帽一族大吃一驚。同時，路飛一行人來到了小島的表面，那裡矗立著未來派建築，還有不知名的巨大生物在遊蕩。

當魯夫等人正在享受蛋頭的未來機器時，一個酷似巴塞洛繆-庫瑪的人追了過來。當魯夫試圖反擊時，邦妮阻止了他，並揭露了她與庫瑪之間令人震驚的關係。

黑鬍子海盜團伏擊了羅帶領的紅心海盜團。黑鬍子海盜團的高層使用各種強大的惡魔果實能力，不斷逼迫紅心海盜團，以獲得路路通副本

沒有魯夫、喬巴和神兵的草帽團被莉莉絲帶到了遠在雲端的實驗室。他們對從未見過的事物感到興奮不已。然而，在實驗室裡，新的威脅正等著他們。

娜美等人遭到了一個酷似神兵衛的帕西菲斯塔人的攻擊，在他們戰鬥的時候，一些維加龐克的衛星在一旁興奮地看著。同時，路飛等人在廢料堆裡發現了一些不尋常的東西。

維加龐克-沙卡與羅賓等人分享了他關於古代王國和百年虛空的理論。他甚至還說，在大原被摧毀後，他還去過大原。與此同時，魯夫和喬巴試圖啟動巨型機器人，令人驚訝的事情發生了。

Shaka reveals the secret behind the foundation of the Revolutionary Army and how the relationship between Vegapunk and Dragon affected it. Meanwhile, Vegapunk explains to Luffy and the others about his special power and how his clones work.

Vegapunk reveals the shocking history of the ancient giant robot and asks Luffy to take him off Egghead. Rob Lucci and his fellow Cipher Pol agents arrive at the island and ask for permission to enter the port.

CP-0 led by Rob Lucci makes a forced landing at Egghead and people on the Fabriophase are plunged into confusion while Shaka and the others prepare to fight back on the Labophase. The enraged Atlas stands up to defend the peace of the island.

Luffy transforms into the Warrior of Liberation as Rob Lucci also shows his awakened Devil Fruit form. As the two clash, Vegapunk reveals the shocking truth of the Gum-Gum Fruit and his eye-opening theory on Devil Fruits.

Vegapunk describes how he managed to grant the Seraphim Devil Fruit powers, which surprises Franky and the others. Meanwhile, Luffy is still facing Lucci in his savage, awakened form, but the rocket to the Labophase is about to leave.

Vegapunk asks the Straw Hats to take him on board their ship which makes Franky and Usopp happy but annoys Nami. Lucci orders his troops to block off all Vegapunk’s escape routes and looks to invade the heavily-secured Labophase.

CP-0 enters the Labophase and starts attacking the Thousand Sunny as soon as they find it. Luffy reunites with Nami and the others but finds out that Vegapunk is missing. Meanwhile, Bonney is pressing Vegapunk to change back her father Kuma.

The Seraphim come following CP-0 to the Labophase, pitting Zoro and Brook against the seven of them. Sanji, Franky, Edison, and Lilith leave the lab to fight them, but the Seraphim launch a full-scale attack on the lab.

Stussy suddenly knocked out Lucci to the Straw Hats’ surprise, and when Shaka reveals her true identity, they’re stunned again. Meanwhile, a major incident occurs on an island called Sphinx where Whitebeard came from.

CP-0 takes control of the Fabriophase but gets attacked by an unknown force. Meanwhile, in the Labophase, the Straw Hats and Dr. Vegapunk’s Satellites find out that Vegapunk has disappeared and begin searching for him.

Bonney witnesses her father Bartholomew Kuma’s painful memories. Meanwhile at the lab, Shaka notices that their communications and surveillance cameras are being cut off by somebody.

The Seraphim suddenly start attacking the Straw Hats and the Vegapunk Satellites. They are not stopping even when the Satellites order them to. When Luffy and Zoro are cornered, Lucci and Kaku wake up and make them a surprising offer.

Lucci and Kaku from CP-0, who came to Egghead with a mission to eliminate the seven Vegapunks, ask Luffy and Zoro to uncuff them so they can fight alongside them against the Seraphim, but Luffy and Zoro are reluctant to do so.

The Straw Hats and CP-0 relentlessly attack the Seraphim, but the Seraphim keep getting back up. Then, Zoro notices that the Seraphim remind him of someone who he fought before. Meanwhile, Shaka goes to check one place where he thinks Vegapunk Stella might be.

Stussy and Sentomaru discuss how big the threat that closes in on Egghead is as Luffy and the others keep fighting the uncontrollable Seraphim. Meanwhile, in an underground lab, someone approaches Vegapunk, who is being held captive.

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