The Cunninghams and Fonzie go on vacation to Lake Pinewood. Richie and Fonzie get together with two lonely milkmaids, and, when their father catches them in his barn with his daughters, he says they will have to get married.

The crossover continues on Laverne & Shirley S05E01 Shotgun Wedding (II).

Wanting to be treated more like an adult, Chachi sells his soul to the devil. In order to get him out of it, Fonzie has to go 24 hours without receiving a kiss from a girl.

The merchant seamen are in town, and with them is Rico, a sworn enemy of Fonzie. Fonzie is primed to fight, but Father Delvecchio makes him promise not to.

Marion forgets to set the parking brake in her car, and it rolls down a hill and through the wall at Arnold's. To show Howard that she is a responsible person, she goes to court to take full responsibility for her actions. She phones Fonzie at Arnold's and says she is in jail.

Wanting to have some money of his own, Richie gets a job loading papers for The Milwaukee Journal. A man named Frank, who works in the same department, doesn't like Richie right from the start.

Angry with Lori Beth about her wanting him to wear a Humpty Dumpty costume for a Homecoming parade, Richie goes to a bar to be alone. He meets a young woman there and finds they work in complementary fields. The woman asks Richie to go with her on a trip to Venice, and he accepts.

Marion is a star in a community theatre play, and her director/co-star begins to make unwelcome advances toward her.

Howard puts on a burlesque show for his Leopard Lodge with the hope that it will earn him the honor of being the Grand Poobah. The show is in jeopardy of being halted when the girls hired for the show are snowed-in in Buffalo, New York.

In a somewhat rebellious move after Howard and Marion won't let her go to Chicago for a concert, Joanie decides to become a model. However, she is unaware that the studio shoots nudes.

Richie is chosen the 1961 Delta Gamma King and acts like a jerk after receiving the honor. However, nobody knows that the reason he acts the way he does is a result of someone slipping a pill into his ginger ale. The next day, everyone is angry with Richie, but he is confused because he doesn't remember anything that happened after being crowned king.

Unknowingly, Richie, Potsie, and Ralph are invited to a party hosted by the She Devils gang in which the Devil who brings in the biggest nerd wins a $100 prize. After Richie discovers the purpose of the party, he and the guys head for the door. However, several Devils come in carrying a sack containing a guy who jilted the sister of the gang's leader. The guy is Chachi.

A wheelchair-bound man, injured by a drunk driver, answers Fonzie's want advertisement for Bronko's Auto Repairing and is hired. He drives Fonzie crazy and is rude to everyone he comes into contact with.

Milwaukee's East Boulevard Business Association, led by Howard, wants to build an expressway off-ramp through Inspiration Point. Everyone does their best to stop the construction and save Inspiration Point.

After initially being reluctant to go through with it, Howard agrees to have a second wedding where he and Marion will renew their vows.

Joanie is unsure about whether or not she wants to go with her boyfriend to The Alamo, a special section of Inspiration Point.

Chachi accidentally sets Arnold's on fire, and Fonzie, Potsie, and Ralph are trapped inside.

As 50/50 partners, Fonzie and Al are at odds about various aspects of the new Arnold's and threaten to not build it at all.

After hearing from a former employee about the wonders of television advertising, Howard thinks it will help his hardware store. He hires someone to handle the advertisement who guarantees a 40 percent increase in business in exchange for complete control of the project.

Fonzie meets a deaf woman at the electric company office where he is attempting to straighten out a billing error for Howard. He gives Richie the indication that this girl may be the one.

Joanie thinks Chachi is losing interest in her, and Chachi thinks Joanie isn't taking him seriously. On the advice of Lori Beth, Joanie invites Chachi over for dinner on a night where she is babysitting Potsie's cousins. Chachi tells Al that he's finally going to make his move with Joanie.

Richie goes to a Leopard Lodge convention with Howard in Chicago despite wanting to go to a Bears game with Potsie and Ralph.

Potsie secures an audition as a singer for a local club. The potential success goes to his head, and he treats those around him as inferiors.

Howard's Uncle Joe, after reading Richie's report on the 1920s, tells him a story about what really happened during Prohibition.

Ralph's parents are getting a divorce, and Ralph becomes depressed.

Meanwhile, Marion practices for her upcomming role as the witch in a Hansel & Gretel play.